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Abel Nightroad, also known as Crusnik 02, is the main protagonist in the series. Nightroad, born Abel Knightlord, is a member of the AX division of the Vatican Ministry of Holy Affairs. As an agent of the AX, he is referred to by his codename, "Crusnik." Originally he hated humans because of their role in the Crusniks' creation and experimentation, but ultimately changed his mind after the death of Lilith Sahl.